Space X-Philes
Astrotolerant Organisms
Introduction
Astrotolerant prokaryotes are defined here as prokaryotes with one or more specific “astrotolerances”, or adaptations/exaptations to enduring interplanetary conditions, such as low temperature, low pressure, high radiation, extreme drouth, and (because interplanetary transit involves vast distances) time. Therefore, included in this category are cryophiles, hypobarophiles, radiotolerant organisms, xerophiles, and chronotolerant organisms. These are in contrast to hypergeophiles, or creatures adapted for surviving planetary extremes.
In addition to other sources cited, much of this information was compiled with the help of resources such as the The List of Prokaryotic names with Standing in Nomenclature (LPSN), and NCBI’s Taxonomy Browser.
Taxonomic/Phylogenetic Overview of Astrotolerant Genera
“Astrotolerant genera” refers here to genera known to include species with astrotolerances.
Genera containing cryophilic and cryotolerant organisms are marked with an asterisk (*)
Genera containing radiophilic and radioresistant organisms are marked with a dagger (†)
Genera containing hypobarophilic and hypobarotolerant organisms are marked with two asterisks (**)
Genera containing xerophilic and xerotolerant organisms are marked with two daggers (††)
Genera containing chronotolerant organisms are marked with a double dagger (‡)
In addition to other sources cited, much of this information was compiled with the help of NCBI’s Taxonomy Browser.
- Domain: Archaea
- Phylum: Euryarchaeota
- Class: Halobacteria
- Order: Halobacteriales
- Family: Halobacteriaceae
- Genus: Halobacterium †+
- Order: Natrialbales
- Family: Natrialbaceae
- Genus: Natrialba ***†††+
- Class: Thermococci
- Order: Thermococcales
- Family: Thermococcaceae
- Genus: Pyrococcus †+
- Genus: Thermococcus †+
- Domain: Bacteria
- Phylum: Actinobacteria
- Class: Rubrobacteria
- Order: Rubrobacterales
- Family: Rubrobacteraceae
- Genus: Rubrobacter †
- Phylum: Deinococcus-Thermus
- Class: Deinococci
- Order: Deinococcales
- Family: Deinococcaceae
- Genus: Deinococcus ***†††+
- Phylum: Proteobacteria
- Class: Alphaproteobacteria
- Order: Hyphomonadales
- Family: Hyphomonadaceae
- Genus: Hyphomonas *
- Class: Betaproteobacteria
- Order: Burkholderiales
- Family: Alcaligenaceae
- Genus: Achromobacter *
- Genus: Alcaligenes *
- Order: Neisseriales
- Family: Chromobacteriaceae
- Genus: Aquaspirillum *
- Class: Gammaproteobacteria
- Order: Alteromonadales
- Family: Alteromonadaceae
- Genus: Alteromonas *
- Family: Colwelliaceae
- Genus: Colwellia *
- Order: Pseudomonadales
- Family: Moraxellaceae
- Genus: Psychrobacter *†
- Kingdom: Fibrobacteres/Chlorobi/Bacteroidetes group
- Superphylum: Bacteroides/Chlorobi group
- Phylum: Bacteroidetes
- Class: Bacteroidia
- Order: Bacteroidales
- Family: Bacteroidaceae
- Genus: Bacteroides *
- Class: Cytophagia
- Order: Cytophagales
- Family: Cytophagaceae
- Genus: Cytophaga *
- Kingdom: Terrabacteria
- Phylum: Actinobacteria
- Class: Actinomycetia
- Order: Micrococcales
- Family: Brevibacteriaceae
- Genus: Brevibacterium *
- Family: Micrococcaceae
- Genus: Arthrobacter *
- Phylum: Firmicutes
- Class: Bacilli
- Order: Bacillales
- Family: Bacillaceae
- Genus: Bacillus ***†††‡+
- Genus: Virgibacillus ***†††‡+
- Family: Incertae sedis
- Genus: Exiguobacterium **
- Class: Clostridia
- Order: Clostridiales
- Family: Clostridiaceae
- Genus: Clostridium ***†††‡+
- Phylum: Cyanobacteria/Melainabacteria group
- Class: Cyanobacteria
- Order: Chroococcidiopsidales
- Family: Chroococcidiopsidaceae
- Genus: Chroococcidiopsis *†+
List of Astrotolerant Genera
“Astrotolerant genera” refers here to genera known to include species with astrotolerances.
Genera containing cryophilic and cryotolerant organisms are marked with an asterisk (*)
Genera containing radiophilic and radioresistant organisms are marked with a dagger (†)
Genera containing hypobarophilic and hypobarotolerant organisms are marked with two asterisks (**)
Genera containing xerophilic and xerotolerant organisms are marked with two daggers (††)
Genera containing chronotolerant organisms are marked with a double dagger (‡)
In addition to other sources cited, much of this information was compiled with the help of NCBI’s Taxonomy Browser.
| Genus | Source |
|---|---|
| * Achromobacter | Morita & Moyer, 2001 |
| * Alcaligenes | Morita & Moyer, 2001 |
| * Alteromonas | Morita & Moyer, 2001 |
| * Aquaspirillum | Morita & Moyer, 2001 |
| * Arthrobacter | Morita & Moyer, 2001 |
| ***†††‡ Bacillus | ? |
| * Bacteroides | Morita & Moyer, 2001 |
| * Brevibacterium | Morita & Moyer, 2001 |
| *** Carnobacterium | ? |
| ***†††‡ Clostridium | ? |
| * Colwellia | Morita & Moyer, 2001 |
| *** Cryobacterium | ? |
| * Cytophaga | Morita & Moyer, 2001 |
| ***††† Deinococcus | ? |
| ** Exiguobacterium | Schuerger & Nicholson, 2016 |
| * Flavobacterium | Morita & Moyer, 2001 |
| * Gelidibacter | Morita & Moyer, 2001 |
| * Halomonas | Wang &al., 2020 |
| * Hyphomonas | Edwards &al., 2003 |
| * Methanococcoides | Morita & Moyer, 2001 |
| * Methanogenium | Morita & Moyer, 2001 |
| * Methanosarcina | Morita & Moyer, 2001 |
| * Methylococcus | Morita & Moyer, 2001 |
| * Microbacterium | Morita & Moyer, 2001 |
| * Micrococcus | Morita & Moyer, 2001 |
| * Moritella | Morita & Moyer, 2001 |
| ***††† Natrialba | (Tindall &al., 1984) Kamekura &al., 1997 |
| * Octadecabacter | Morita & Moyer, 2001 |
| ** Paenibacillus | ? |
| * Phormidium | Morita & Moyer, 2001 |
| * Photobacterium | Morita & Moyer, 2001 |
| * Polaribacter | Morita & Moyer, 2001 |
| * Polaromonas | Morita & Moyer, 2001 |
| * Psuedomonas | Morita &al., 2001; Saurabh &al., 2020; Yumoto &al., 2001 |
| *† Psychrobacter | Garcia-Lopez &al., 2014 |
| * Psychroserpens | Morita & Moyer, 2001 |
| ** Rhodococcus | ? |
| † Rubrobacter | Kampfer &al., 2014 |
| * Shewanella | Morita & Moyer, 2001 |
| ** Streptomyces | ? |
| ** Serratia | ? |
| † Thermococcus | ? |
| * Vibrio | Morita & Moyer, 2001 |
List of Astrotolerant Species
In addition to other sources cited, much of this information was compiled with the help of resources such as the The List of Prokaryotic names with Standing in Nomenclature (LPSN), and NCBI’s Taxonomy Browser.
Note: This list is still under construction and has definitely not yet been double-checked for accuracy! This situation will be remedied as quickly as possible.
| Genus & Species | Source |
|---|---|
| * Achromobacter aegrifaciens | Vandamme &al., 2014 |
| * Achromobacter agilis | Vandamme &al., 2016 |
| * Achromobacter aloeverae | Kuncharoen &al., 2017 |
| * Achromobacter animicus | Vandamme &al., 2013 |
| * Achromobacter anxifer | Vandamme &al., 2014 |
| * Achromobacter deleyi | Vandamme &al., 2016 |
| * Achromobacter dentrificans | (Ruger &Tan, 1983) Coenye &al., 2003 |
| * Achromobacter dolens | Vandamme &al., 2016 |
| * Achromobacter insolitus | Coenye &al., 2003 |
| * Achromobacter insuavis | Vandamme &al., 2014 |
| * Achromobacter kerstersii | Vandamme &al., 2016 |
| * Achromobacter marplatensis | Gomila &al., 2011 |
| * Achromobacter mucicolens | Vandamme &al., 2013 |
| * Achromobacter pestifer | (Frankland & Frankland, 1887) Vandamme &al., 2016 |
| * Achromobacter piechaudii | (Kiredjian &al., 1986) Yabuuchi &al., 1998 |
| * Achromobacter pulmonis | Vandamme &al., 2013 |
| * Achromobacter ruhlandii | (Packer &Vishniac, 1955) Yabuuchi &al., 1998 |
| * Achromobacter sediminum | Zhang &al., 2014 |
| * Achromobacter spanius | Coenye &al., 2003 |
| * Achromobacter spiritinus | Vandamme &al., 2013 |
| * Achromobacter veterisilvae | Dumolin &al., 2020 |
| * Achromobacter xylosoxidans | (Yabuuchi & Ohyama, 1971) Yabuuchi & Yano, 1981 |
| * Achromobacter ssp. | Morita & Moyer, 2001 |
| * Alcaligenes aquatilis | Van Trappen &al., 2005 |
| * Alcaligenes endophyticus | Lu &al., 2017 |
| * Alcaligenes faecalis | Castellani & Chalmers, 1919 |
| * Alcaligenes pakistanensis | Abbas &al., 2016 |
| * Alcaligenes ssp. | Morita & Moyer, 2001 |
| * Alteromonas addita | Ivanova &al., 2005 |
| * Alteromonas aestuariivivens | Park &al., 2017 |
| * Alteromonas alba | Sun &al., 2019 |
| * Alteromonas australica | Ivanova &al., 2013 |
| * Alteromonas confluentis | Park &al., 2015 |
| * Alteromonas facilis | Zhang &al., 2020 |
| * Alteromonas flava | Zhang &al., 2020 |
| * Alteromonas fortis | Barbeyron &al., 2019 |
| * Alteromonas genovensis | Vandecandelaere &al., 2008 |
| * Alteromonas gracilis | Matsuyama &al., 2015 |
| *+ Alteromonas halophila (halophile?) | Chen &al., 2012 |
| * Alteromonas hispanica | Martinez-Checa &al., 2005 |
| * Alteromonas lipolytica | Shi &al., 2017 |
| * Alteromonas litorea | Yoon &al., 2003 |
| * Alteromonas macleodii | Baumann &al., 1972 |
| * Alteromonas marina | Yoon &al., 2003 |
| * Alteromonas mediterranea | Ivanova &al., 2015 |
| * Alteromonas naphthalenivorans | Jin &al., 2015 |
| * Alteromonas nigrifaciens | (White &al., 1940) Baumann &al., 1984 |
| * Alteromonas oceani | Jin &al., 2018 |
| * Alteromonas pelagimontana | Sinha &al., 2017 |
| * Alteromonas marina | Yoon &al., 2003 |
| * Alteromonas ponticola | Kim &al., 2020 |
| * Alteromonas portus | Huang &al., 2020 |
| * Alteromonas profundi | Shen &al., 2020 |
| * Alteromonas sediminis | Ye &al., 2019 |
| * Alteromonas simiduii | Chiu &al., 2007 |
| * Alteromonas stellipolaris | Van Trappen &al., 2004 |
| * Alteromonas tagae | Simidu &al., 1990 |
| * Alteromonas ssp. | Morita & Moyer, 2001 |
| * Aquaspirillum arcticum | Butler &al., 1990 |
| * Aquaspirillum bengal | Kumar &al., 1974 |
| * Aquaspirillum polymorphum | (Williams & Ritten, 1980) Hylemon &al., 1973 |
| * Aquaspirillum putridiconchylium | (Terasaki, 1961) Hylemon &al., 1973 |
| * Aquaspirillum serpens | (Muller, 1786) Hylemon &al., 1973 |
| * Aquaspirillum soli | Moya &al., 2017 |
| * Aquaspirillum ssp. | Morita & Moyer, 2001 |
| * Arthrobacter agilis | (Ali-Cohen, 1889) Koch &al., 1995 |
| *+ Arthrobacter alkaliphilus | Ding &al., 2009 |
| * Arthrobacter alpinus | Zhang &al., 2010 |
| * Arthrobacter bambusae | Park &al., 2014 |
| * Arthrobacter bussei | Flegler &al., 2020 |
| * Arthrobacter castelli | Heyrman &al., 2005 |
| * Arthrobacter celericrescens | Yan &al., 2019 |
| * Arthrobacter citreus | Sacks, 1954 |
| * Arthrobacter cryoconiti | Margesin &al., 2012 |
| * Arthrobacter crystallopoietes | Ensign & Rittenberg, 1963 |
| * Arthrobacter cupressi | Zhang &al., 2012 |
| * Arthrobacter deserti | Hu &al., 2016 |
| * Arthrobacter echini | Lee &al., 2016 |
| * Arthrobacter flavus | Reddy &al., 2000 |
| * Arthrobacter gandavensis | Storms &al., 2003 |
| * Arthrobacter ginkgonis | Cheng &al., 2017 |
| * Arthrobacter glacialis | Liu &al., 2019 |
| * Arthrobacter globiformis | (Conn, 1928) Conn & Dimmick, 1947 |
| * Arthrobacter gyeryongensis | Hoang &al., 2014 |
| *+? Arthrobacter halodurans | Chen &al., 2012 |
| * Arthrobacter humicola | Kageyama &al., 2008 |
| * Arthrobacter koreensis | Lee &al., 2003 |
| * Arthrobacter liuii | Yu &al., 2015 |
| * Arthrobacter livingstonensis | Ganzert &al., 2011 |
| * Arthrobacter luteolus | Wauters &al., 2000 |
| * Arthrobacter methylotrophus | Borodina &al., 2002 |
| * Arthrobacter mobilis | Ye &al., 2020 |
| * Arthrobacter monumenti | Heyrman &al., 2005 |
| * Arthrobacter nanjingensis | Huang &al., 2015 |
| * Arthrobacter orzyae | Kageyama &al., 2008 |
| * Arthrobacter paludis | Zhang &al., 2018 |
| * Arthrobacter parietis | Heyrman &al., 2005 |
| * Arthrobacter pascens | Lochhead & Burton, 1953 |
| * Arthrobacter pigmenti | Heyrman &al., 2005 |
| * Arthrobacter pityocampae | Ince &al., 2014 |
| * Arthrobacter pokkalii | Krishnan &al., 2017 |
| * Arthrobacter psychrochitiniphilus | Wang &al., 2009 |
| * Arthrobacter psychrolactophilus | Loveland-Curtze &al., 2000 |
| * Arthrobacter ramosus | Jensen, 1960 |
| * Arthrobacter rhombi | Osorio &al., 1999 |
| * Arthrobacter roseus | Reddy &al., 2002 |
| * Arthrobacter ruber | Liu &al., 2018 |
| * Arthrobacter russicus | Li &al., 2004 |
| * Arthrobacter silvisoli | Yan &al., 2018 |
| * Arthrobacter silviterrae | Lee &al., 2017 |
| * Arthrobacter stackerbrandtii | Tvrzova &al., 2005 |
| * Arthrobacter subterraneus | Chang &al., 2008 |
| * Arthrobacter tecti | Heyrman &al., 2005 |
| * Arthrobacter terricola | Trinh & Kim, 2021 |
| * Arthrobacter tumbae | Heyrman &al., 2005 |
| * Arthrobacter ulcerisalmonis | Kampfer &al., 2020 |
| * Arthrobacter woluwensis | Funke &al., 1997 |
| * Arthrobacter yangruifuii | Ge &al., 2020 |
| * Arthrobacter zhaeoguopingii | Ge &al., 2020 |
| * Arthrobacter ssp. | Morita & Moyer, 2001 |
| ***†††‡ Bacillus acidicola | Albert &al. 2005 |
| ***†††‡ Bacillus aerius | Shivaji &al., 2014 |
| ***†††‡ Bacillus aerophilus | Shivaji &al., 2006 |
| ***†††‡ Bacillus albus | Liu &al., 2017 |
| ***†††‡ Bacillus altitudinis | Shivaji &al., 2006 |
| ***†††‡ Bacillus alveayuensis | Bae &al., 2005 |
| ***†††‡ Bacillus amyloliquefaciens | (Fukumoto, 1943) Priest &al., 1987 |
| ***†††‡ Bacillus anthracis (Anthrax) | Cohn, 1872 |
| ***†††‡ Bacillus aquiflavi | Xie &al., 2020 |
| ***†††‡ Bacillus atrophaeus | Nakamura, 1989 |
| ***†††‡ Bacillus australimaris | Liu &al., 2016 |
| ***†††‡ Bacillus badius | Batchelor, 1919 |
| ***†††‡ Bacillus benzoevorans | Pichinoty &al., 1987 |
| ***†††‡ Bacillus cabrialesii | de Los Santos Villalobos &al., 2019 |
| ***†††‡ Bacillus canaveralius | Newcombe &al., 2009 |
| ***†††‡ Bacillus capparidis | Wang &al., 2017 |
| ***†††‡ Bacillus cereus | Frankland & Frankland, 1887 |
| ***†††‡ Bacillus chungangensis | Cho &al., 2010 |
| ***†††‡ Bacillus cahuilensisereus | Cerritos &al., 2008 |
| ***†††‡ Bacillus cytotoxicus | Guinebretiere &al., 2013 |
| ***†††‡ Bacillus decisifrondis | Zhang &al., 2007 |
| ***†††‡ Bacillus ectoiniformans | Zhu &al., 2016 |
| ***†††‡ Bacillus enclensis | Dastager &al., 2014 |
| ***†††‡ Bacillus fengquiuensis | Zhao &al., 2014 |
| ***†††‡ Bacillus fungorum | Liu &al., 2020 |
| ***†††‡ Bacillus glycinifermentans | Kim &al., 2015 |
| ***†††‡ Bacillus gobiensisectoiniformans | Liu &al., 2016 |
| ***†††‡ Bacillus halotolerans | (Delaporte & Sasson, 1967) Tindall &al., 2017 |
| ***†††‡ Bacillus heynesii | Dunlap &al., 2017 |
| ***†††‡ Bacillus horti | Yumoto &al., 1998 |
| ***†††‡ Bacillus inaquosorum | (Rooney &al., 2009) Dunlap &al., 2020 |
| ***†††‡ Bacillus infantis | Ko &al., 2006 |
| ***†††‡ Bacillus infernus | Boone &al., 1995 |
| ***†††‡ Bacillus isabeliae | Albuquerque &al., 2008 |
| ***†††‡ Bacillus kexueae | Sun &al., 2018 |
| ***†††‡ Bacillus licheniformis | (Weigmann, 1898) Chester, 1901 |
| ***†††‡ Bacillus luti | Liu &al., 2017 |
| ***†††‡ Bacillus manusensis | Sun &al., 2018 |
| ***†††‡ Bacillus marinisedimentorum | Guo &al., 2018 |
| ***†††‡ Bacillus mesophilus | Manickam &al., 2014 |
| ***†††‡ Bacillus methanolicus | Arfman &al., 1992 |
| ***†††‡ Bacillus mobilis | Liu &al., 2017 |
| ***†††‡ Bacillus mojavensis | Roberts &al., 1994 |
| ***†††‡ Bacillus mycoides | Flugge &al., 1886 |
| ***†††‡ Bacillus nakamurai | Dunlap &al., 2016 |
| ***†††‡ Bacillus ndiopicus | Lo &al., 2017 |
| ***†††‡ Bacillus nitratireducens | Guo &al., 2016 |
| ***†††‡ Bacillus oleivorans | Azmatunnisa &al., 2015 |
| ***†††‡ Bacillus pacificus | Liu &al., 2017 |
| ***†††‡ Bacillus pakistanensis | Roohi &al., 2014 |
| ***†††‡ Bacillus paralicheniformis | Dunlap &al., 2015 |
| ***†††‡ Bacillus paramycoides | Liu &al., 2017 |
| ***†††‡ Bacillus paranthracis | Liu &al., 2017 |
| ***†††‡ Bacillus pervagus | Kosowski &al., 2014 |
| ***†††‡ Bacillus priscicola | Lee &al., 2016 |
| ***†††‡ Bacillus proteolyticus | Liu &al., 2017 |
| ***†††‡ Bacillus pseudomycoides | Nakamura, 2018 |
| ***†††‡ Bacillus pumilus | Meyer & Gottheil, 1901 |
| ***†††‡ Bacillus safensis | Satomi &al., 2006 |
| ***†††‡ Bacillus salacetis | Daroonpunt &al., 2019 |
| ***†††‡ Bacillus salinus | Kang &al., 2020 |
| ***†††‡ Bacillus salitolerans | Zhang &al., 2015 |
| ***†††‡ Bacillus seohaeanensis | Lee &al., 2006 |
| ***†††‡ Bacillus siamensis | Sumpavapol &al., 2010 |
| ***†††‡ Bacillus shivajii | Kumar &al., 2018 |
| ***†††‡ Bacillus smithii | Nakamura &al., 1988 |
| ***†††‡ Bacillus solimangrovi | Lee &al., 2014 |
| ***†††‡ Bacillus songklensis | Kang &al., 2013 |
| ***†††‡ Bacillus sonoresnis | Palmisano &al., 2001 |
| ***†††‡ Bacillus spizizenii | (Nakamura &al., 1999) Dunlap &al., 2020 |
| ***†††‡ Bacillus spongiae | Lee &al., 2019 |
| ***†††‡ Bacillus stercoris | (Adelskov & Patel, 2017) Dunlap &al., 2020 |
| ***†††‡ Bacillus stratosphericus | Shivaji &al., 2006 |
| ***†††‡ Bacillus subtilis | (Ehrenberg 1835) Cohn 1872 |
| ***†††‡ Bacillus swezeyi | Dunlap &al., 2017 |
| ***†††‡ Bacillus taeanensis | Lim &al., 2006 |
| ***†††‡ Bacillus tamaricis | Zhang &al., 2018 |
| ***†††‡ Bacillus tequilensis | Gatson &al., 2006 |
| ***†††‡ Bacillus thermocloacae | Demharter & Hensel, 1989 |
| ***†††‡ Bacillus thermotolerans | Yang &al., 2013 |
| ***†††‡ Bacillus thuringiensis | Burlinger, 1915 |
| ***†††‡ Bacillus tianshenii | Jiang &al., 2014 |
| ***†††‡ Bacillus toyonensis | Jimenez &al., 2014 |
| ***†††‡ Bacillus tropicus | Liu &al., 2017 |
| ***†††‡ Bacillus vallismortis | Roberts &al., 1996 |
| ***†††‡ Bacillus velezensis | Ruis-Garcia &al., 2005 |
| ***†††‡ Bacillus wiedmannii | Miller &al., 2016 |
| ***†††‡ Bacillus wudalienchiensis | Liu &al., 2017 |
| ***†††‡ Bacillus xiamenensis | Lai &al., 2014 |
| ***†††‡ Bacillus xiapuensis | Liu &al., 2019 |
| ***†††‡ Bacillus zhangzhouensis | Liu &al., 2016 |
| ***†††‡ Bacillus sp. | Schuerger & Nicholson, 2016 |
| * Bacteroides acidifaciens | Miyamoto & Itoh, 2000 |
| * Bacteroides caccae | Johnson &al., 1986 |
| * Bacteroides caecicola | Irisawa &al., 2016 |
| * Bacteroides caecigallinarum | Saptura &al., 2015 |
| * Bacteroides caecimuris | Lagkouvardos &al., 2016 |
| * Bacteroides cellulosilyticus | Robert &al., 2007 |
| * Bacteroides clarus | Watanabe &al., 2010 |
| * Bacteroides coprosuis | Whitehead &al., 2005 |
| * Bacteroides eggerthii | Holdeman & Moore, 1974 |
| * Bacteroides faecalis | Yu &al., 2019 |
| * Bacteroides faecichinchillae | Kitahara &al., 2012 |
| * Bacteroides faecis | Kim &al., 2010 |
| * Bacteroides finegoldii | Bakir &al., 2006 |
| * Bacteroides fluxus | Watanabe &al., 2010 |
| * Bacteroides fragilis | (Veillon & Zuber, 1898) Castellani & Chalmers, 1919 |
| * Bacteroides galacturonicus | Jensen & Canale-Parola, 1987 |
| * Bacteroides gallinaceum | Irisawa &al., 2016 |
| * Bacteroides gallinarum | Lan &al., 2006 |
| * Bacteroides graminisolvens | Nishiyama &al., 2009 |
| * Bacteroides helcogenes | Benno &al., 1983 |
| * Bacteroides instestinalis | Bakir &al., 2006 |
| * Bacteroides koreensis | Shin &al., 2017 |
| * Bacteroides kribbi | Shin &al., 2017 |
| * Bacteroides luti | Hatamoto &al., 2014 |
| * Bacteroides nordii | Song &al., 2005 |
| * Bacteroides oleiciplenus | Watanabe &al., 2010 |
| * Bacteroides ovatus | Eggerth & Gagnon, 1933 |
| * Bacteroides pectinophilus | Jensen & Canale-Parola, 1987 |
| * Bacteroides polypragmatus | Patel & Breuil, 1982 |
| * Bacteroides propionicifaciens | Ueki &al., 2008 |
| * Bacteroides pyrogenes | Benno &al., 1983 |
| * Bacteroides reticulotermitis | Sakamoto & Ohkuma, 2013 |
| * Bacteroides rodentium | Kitahara &al., 2011 |
| * Bacteroides salyersiae | Song &al., 2005 |
| * Bacteroides stercorirosoris | Kitahara &al., 2012 |
| * Bacteroides stercoris | Johnson &al., 1986 |
| * Bacteroides thetataiotaomicron | (Distaso, 1912) Castellani & Chalmers, 1919 |
| * Bacteroides uniformis | Eggerth & Gagnon, 1933 |
| * Bacteroides xylanolyticus | Scholten-Koerselman &al., 1988 |
| * Bacteroides ssp. | Morita & Moyer, 2001 |
| * Brevibacterium album | Tang &al., 2008 |
| * Brevibacterium ammoniilyticum | Kim &al., 2013 |
| * Brevibacterium anseongense | Jung &al., 2019 |
| * Brevibacterium antiquum | Gavrish &al., 2005 |
| * Brevibacterium aurantiacum | Gavrish &al., 2005 |
| * Brevibacterium avium | Pascual & Collins, 1999 |
| * Brevibacterium casei | Collins &al., 1983 |
| * Brevibacterium celere | Ivanova &al., 2004 |
| * Brevibacterium daeguense | Cui &al., 2013 |
| * Brevibacterium epidermidis | Collins &al., 1983 |
| * Brevibacterium frigoritolerans | Delaporte & Sasson, 1967 |
| * Brevibacterium hankyongi | Choi &al., 2018 |
| * Brevibacterium iodinum | (Davis, 1939) Collins &al., 1981 |
| * Brevibacterium jeotgali | Choi &al., 2013 |
| * Brevibacterium linens | (Wolff, 1910) Breed, 1953 |
| * Brevibacterium luteolum | Wauters &al., 2003 |
| * Brevibacterium linens | (Wolff, 1910) Breed, 1953 |
| * Brevibacterium marinum | Lee, 2008 |
| * Brevibacterium mcbrellneri | McBride &al., 1994 |
| * Brevibacterium oceani | Bhadra &al., 2008 |
| * Brevibacterium otitidis | Pascual, 1996 |
| * Brevibacterium paucivorans | Wauters &al., 2001 |
| * Brevibacterium permense | Gavrish &al., 2005 |
| * Brevibacterium picturae | Heyrman &al., 2004 |
| * Brevibacterium pityocampae | Kati &al., 2010 |
| * Brevibacterium profundi | Pei &al., 2020 |
| * Brevibacterium ravenspurgense | Mages &al., 2009 |
| * Brevibacterium rongguiense | Deng &al., 2020 |
| *+ Brevibacterium salitolerans | Guan &al., 2010 |
| * Brevibacterium samyangense | Lee, 2006 |
| * Brevibacterium sandarakinum | Kampfer &al., 2010 |
| * Brevibacterium sanguinis | Wauters &al., 2004 |
| * Brevibacterium sediminis | Chen &al., 2004 |
| * Brevibacterium senegalense | Kokcha &al., 2013 |
| * Brevibacterium siliguriense | Kumar &al., 2013 |
| * Brevibacterium yomogidense | Tonouchi &al., 2013 |
| * Brevibacterium ssp. | Morita & Moyer, 2001 |
| *** Carnobacterium alterfunditum | Nicholson &al., 2013 |
| *** Carnobacterium divergens | Nicholson &al., 2013 |
| *** Carnobacterium funditum | Nicholson &al., 2013 |
| *** Carnobacterium gallinarum | Nicholson &al., 2013 |
| *** Carnobacterium inhibens | Nicholson &al., 2013 |
| *** Carnobacterium maltaromaticum | Nicholson &al., 2013 |
| *** Carnobacterium mobile | Nicholson &al., 2013 |
| *** Carnobacterium pleistocenium | Nicholson &al., 2013 |
| *** Carnobacterium viridans | Nicholson &al., 2013 |
| *** Carnobacterium sp. | Schuerger & Nicholson, 2016 |
| ***†††‡ Clostridium aceticum | Wieringa, 1940 |
| ***†††‡ Clostridium acetireducens | Orlygsson &al., 1996 |
| ***†††‡ Clostridium acetobutylicum | McCoy &al., 1926 |
| ***†††‡ Clostridium acidisoli | Kuhner &al., 2000 |
| ***†††‡ Clostridium aestuarii | Kim &al., 2007 |
| ***†††‡ Clostridium akagii | Kuhner &al., 2000 |
| ***†††‡ Clostridium algidicarnis | Lawson &al., 1995 |
| ***†††‡ Clostridium algifaecis | Wu &al., 2014 |
| ***†††‡ Clostridium algoriphilum | Shcherbakova &al., 2010 |
| ***†††‡ Clostridium amazonense | O’Neal &al., 2015 |
| ***†††‡ Clostridium aminophilum | Paster &al., 1993 |
| ***†††‡ Clostridium amylolyticum | Song & Dong, 2008 |
| ***†††‡ Clostridium arbusti | Jung &al., 2010 |
| ***†††‡ Clostridium arcticum | Jordan & McNicol, 1979 |
| ***†††‡ Clostridium argentinense | Suen &al., 1988 |
| ***†††‡ Clostridium atrophaeus | Nakamura, 1989 |
| ***†††‡ Clostridium aurantibutyricum | Hellinger, 1944 |
| ***†††‡ Clostridium baratii | Prevot, 1938 |
| ***†††‡ Clostridium beihaiense | Dong &al., 2018 |
| ***†††‡ Clostridium beijerinckii | Donker, 1926 |
| ***†††‡ Clostridium bornimense | Hahnke &al., 2014 |
| ***†††‡ Clostridium botulinum | van Ermengem, 1896 |
| ***†††‡ Clostridium bovifaecis | Zhu &al., 2018 |
| ***†††‡ Clostridium bowmanii | Spring &al., 2003 |
| ***†††‡ Clostridium budayi | Le Blaye, 1914 |
| ***†††‡ Clostridium butyricum | Prazmowski, 1880 |
| ***†††‡ Clostridium cadaveris | Klein, 1899 |
| ***†††‡ Clostridium carboxidivorans | Liou &al., 2005 |
| ***†††‡ Clostridium carnis | Klein, 1904 |
| ***†††‡ Clostridium cavendishii | Bowman &al., 2010 |
| ***†††‡ Clostridium celatum | Hauschild & Holdeman, 1974 |
| ***†††‡ Clostridium cellulosi | He &al., 1991 |
| ***†††‡ Clostridium cellulovorans | Sleat &al., 1985 |
| ***†††‡ Clostridium chartatabidum | Kelly &al., 1996 |
| ***†††‡ Clostridium chauvoei | Arloing &al., 1887 |
| ***†††‡ Clostridium chromiireducens | Inglett, 2011 |
| ***†††‡ Clostridium clostridioforme | Burri & Ankersmit, 1906 |
| ***†††‡ Clostridium cochlearium | Douglas, 1919 |
| ***†††‡ Clostridium cocleatum | Kaneuchi &al., 1979 |
| ***†††‡ Clostridium colicanis | Greetham &al., 2003 |
| ***†††‡ Clostridium colinum | Berkhoff &al., 1974 |
| ***†††‡ Clostridium collagenovorans | Jain & Zeikus, 1988 |
| ***†††‡ Clostridium combesii | Prevot & Laplanche, 1947 |
| ***†††‡ Clostridium composti | Shin &al., 2018 |
| ***†††‡ Clostridium cylindrosporum | Barker & Beck, 1942 |
| ***†††‡ Clostridium disporicum | Horn, 1987 |
| ***†††‡ Clostridium drakei | Liou &al., 2005 |
| ***†††‡ Clostridium estertheticum | Collins &al., 1993 |
| ***†††‡ Clostridium fallax | Weinberg & Seguin, 1915 |
| ***†††‡ Clostridium felsineum | Carbone & Tombolato, 1917 |
| ***†††‡ Clostridium fermenticellae | Xu &al., 2019 |
| ***†††‡ Clostridium fessum | Seo &al., 2020 |
| ***†††‡ Clostridium fimetarium | Kotsyurbenko &al., 1997 |
| ***†††‡ Clostridium formicaceticum | Andreesen &al., 1970 |
| ***†††‡ Clostridium frigidicarnus | Broda &al., 1999 |
| ***†††‡ Clostridium frigoris | Spring &al., 2003 |
| ***†††‡ Clostridium ganghwense | Kim &al., 2006 |
| ***†††‡ Clostridium gasigenes | Broda &al., 2000 |
| ***†††‡ Clostridium granti | Mountfort &al., 1996 |
| ***†††‡ Clostridium guangxiense | Zhao &al., 2017 |
| ***†††‡ Clostridium haemolyticum | Hall, 1929 |
| ***†††‡ Clostridium halophilum | Fendrich &al., 1991 |
| ***†††‡ Clostridium herbivorans | Varel &al., 1995 |
| ***†††‡ Clostridium hiranonis | Kitahara &al., 2001 |
| ***†††‡ Clostridium histolyticum
(not very thermotolerant; OGT=60°C) |
Weinberg & Seguin, 1916 |
| ***†††‡ Clostridium homopropionicum | Dorner & Schink, 1991 |
| ***†††‡ Clostridium huakuii | Ruan &al., 2014 |
| ***†††‡ Clostridium hydrogeniformans | Bowman &al., 2010 |
| ***†††‡ Clostridium hylemonae | Kitahara &al., 2000 |
| ***†††‡ Clostridium indicum
(extremotolerant mesophile, OGT=37°C) |
Gundawar &al., 2019 |
| ***†††‡ Clostridium innocuum | Smith & King, 1962 |
| ***†††‡ Clostridium intestinale | Lee &al., 1989 |
| ***†††‡ Clostridium irregulare | Choukevitch, 1911 |
| ***†††‡ Clostridium isatidis | Padden &al., 1999 |
| ***†††‡ Clostridium jeddahense | Lagier &al., 2016 |
| ***†††‡ Clostridium kluyveri | Barker & Taha, 1942 |
| ***†††‡ Clostridium lacusfryxellense | Spring &al., 2003 |
| ***†††‡ Clostridium leptum | Moore &al., 1976 |
| ***†††‡ Clostridium liquoris | Yin &al., 2016 |
| ***†††‡ Clostridium ljungdahlii | Tanner &al., 1993 |
| ***†††‡ Clostridium lundense | Cirne &al., 2006 |
| ***†††‡ Clostridium luticellarii | Wang &al., 2015 |
| ***†††‡ Clostridium magnum | Schink, 1984 |
| ***†††‡ Clostridium malenominatum | Weinberg &al., 1937 |
| ***†††‡ Clostridium methoxybenzovorans | Mechichi &al., 1999 |
| ***†††‡ Clostridium methylpentosum | Himelbloom & Canale-Parola, 1989 |
| ***†††‡ Clostridium moniliforme | Repaci, 1910 |
| ***†††‡ Clostridium neonatale | Bernard &al., 2018 |
| ***†††‡ Clostridium neuense | Zhao &al., 2017 |
| ***†††‡ Clostridium nexile | Holdeman & Moore, 1974 |
| ***†††‡ Clostridium nitritogenes | Prevot, 1940 |
| ***†††‡ Clostridium nitrophenolicum | Suresh &al., 2007 |
| ***†††‡ Clostridium novyi | Migula, 1900 |
| ***†††‡ Clostridium oceanicum | Smith, 1970 |
| ***†††‡ Clostridium oryzae | Horino &al., 2015 |
| ***†††‡ Clostridium pabulibutyricum | Kobayashi &al., 2017 |
| ***†††‡ Clostridium paradoxum | Li &al., 1993 |
| ***†††‡ Clostridium paraputrificum | Bienstock, 1906 |
| ***†††‡ Clostridium pascui | Wilde &al., 1997 |
| ***†††‡ Clostridium pasteurianum | Winogradsky, 1985 |
| ***†††‡ Clostridium peptidovorans | Mechichi &al., 2000 |
| ***†††‡ Clostridium perfringens
(formerly C. welchii) |
Veillon & Zuber, 1898 |
| ***†††‡ Clostridium phytofermentans | Warnick &al., 2002 |
| ***†††‡ Clostridium piliforme | Tyzzer, 1917 |
| ***†††‡ Clostridium polyendosporum | Duda &al., 1996 |
| ***†††‡ Clostridium polysaccharolyticum | van Glyswyk, 1981 |
| ***†††‡ Clostridium populeti | Sleat & Mah, 1985 |
| ***†††‡ Clostridium prolinivorans | Huang &al., 2020 |
| ***†††‡ Clostridium psychrophilum | Spring &al., 2003 |
| ***†††‡ Clostridium punense | Lanjekar &al., 2015 |
| ***†††‡ Clostridium puniceam | Lund &al., 1981 |
| ***†††‡ Clostridium putrefaciens | McBryde, 1911 |
| ***†††‡ Clostridium quinii | Svensson &al., 1995 |
| ***†††‡ Clostridium ramosum | Veillon & Zuber, 1898 |
| ***†††‡ Clostridium roseum | McCoy & McClung, 1935 |
| ***†††‡ Clostridium saccharobutylicum | Kleis &al., 2001 |
| ***†††‡ Clostridium saccharogumia | Clavel &al., 2007 |
| ***†††‡ Clostridium saccharoperbutylacetonicum | Kleis &al., 2001 |
| ***†††‡ Clostridium sardiniense
(aka Clostridium absonum |
Prevot, 1938 |
| ***†††‡ Clostridium sartagoforme | Partansky & Henry, 1935 |
| ***†††‡ Clostridium saudiense | Angelakis &al., 2016 |
| ***†††‡ Clostridium scidens | Morris &al., 1985 |
| ***†††‡ Clostridium senagelense | Mishra &al., 2015 |
| ***†††‡ Clostridium septicum | Macé, 1889 |
| ***†††‡ Clostridium spiroforme | Kaneuchi &al., 1979 |
| ***†††‡ Clostridium sporogenes | Metchnikoff, 1908 |
| ***†††‡ Clostridium sporosphaeroides | Soriano & Soriano, 1948 |
| ***†††‡ Clostridium subterminale | Hall & Whitehead, 1927 |
| ***†††‡ Clostridium sulfidigenes | Sallam & Steinbuchel, 2009 |
| ***†††‡ Clostridium swellfunianum | Liu &al., 2015 |
| ***†††‡ Clostridium symbiosum | Stevens, 1956 |
| ***†††‡ Clostridium tagluense | Suetin &al., 2009 |
| ***†††‡ Clostridium tarantellae | (Udey &al., 1977) Lawson & Rainey, 2016 |
| ***†††‡ Clostridium tepidiprofundi | Slobodkina &al., 2008 |
| ***†††‡ Clostridium tepidum | Dobritsa &al., 2017 |
| ***†††‡ Clostridium tertium | (Henry, 1917) Bergey &al., 1923 |
| ***†††‡ Clostridium tetani | (Flugge, 1886) Bergey &al., 1923 |
| ***†††‡ Clostridium tetanomorphum | (Bulloch &al., 1919) Wilde &al., 1989 |
| ***†††‡ Clostridium thermarum | Liu &al., 2021 |
| ***†††‡ Clostridium thermoalcaliphilum | Li &al., 1994 |
| ***†††‡ Clostridium thermobutyricum | Weigel &al., 1989 |
| ***†††‡ Clostridium thermopalmarium | Soh &al., 1991 |
| ***†††‡ Clostridium thermopapyrolyticum | Mendez &al., 1991 |
| ***†††‡ Clostridium thermosaccharolyticum | McClung, 1935 |
| ***†††‡ Clostridium thermosuccinogenes | Drent &al., 1995 |
| ***†††‡ Clostridium thermosulfurigenes | Schink & Zeikus, 1983 |
| ***†††‡ Clostridium thiosulfatireducens | Hernandez-Euginio &al., 2002 |
| ***†††‡ Clostridium tyrobutyricum | van Beynum & Pette, 1935 |
| ***†††‡ Clostridium uliginosum | Matthies &al., 2001 |
| ***†††‡ Clostridium villosum | Love &al., 1979 |
| ***†††‡ Clostridium vincentii | Mountfort &al., 1997 |
| ***†††‡ Clostridium viride | Buckel &al., 1995 |
| ***†††‡ Clostridium vulturis | Paek &al., 2015 |
| ***†††‡ Clostridium sp. | Schuerger & Nicholson, 2016 |
| * Colwellia aestuarii | Jung &al., 2006 |
| * Colwellia agarivorans | Xu &al., 2017 |
| * Colwellia aquaemaris | Liu &al., 2014 |
| * Colwellia arctica | Wang &al., 2001 |
| * Candidatus Colwellia aromaticivorans | Campeao &al., 2019 |
| * Colwellia asteriadis | Choi &al., 2010 |
| * Colwellia beringensis | Zhang &al., 2017 |
| * Colwellia chukchiensis | Yu &al., 2011 |
| * Colwellia demingiae | Bowman &al., 1998 |
| * Colwellia echini | Christiansen &al., 2018 |
| * Colwellia hadaliensis | Deming &al., 1988 |
| * Colwellia hornerae | Bowman &al., 1998 |
| * Colwellia marinimaniae | Kusube &al., 2017 |
| * Colwellia maris | Yumoto &al., 1998 |
| * Colwellia meonggei | Kim &al., 2017 |
| * Colwellia mytili | Kim &al., 2017 |
| * Colwellia piezophila | Nogi &al., 2004 |
| * Colwellia polaris | Zhang &al., 2008 |
| * Colwellia ponticola | Park &al., 2019 |
| * Colwellia psychrerythraea | (D’Aoust & Kushner, 1972) Deming &al., 1998 |
| * Colwellia psychotropica | Morita & Moyer, 2001 |
| * Colwellia rossensis | Bowman &al., 1998 |
| * Colwellia sediminilitoris | Park &al., 2016 |
| * Colwellia ssp. | Morita & Moyer, 2001 |
| * Cryobacterium arcticum | Bajerski &al., 2011 |
| * Cryobacterium aureum | Liu &al., 2018 |
| * Cryobacterium flavum | Liu &al., 2012 |
| * Cryobacterium levicorallinum | Liu &al., 2013 |
| * Cryobacterium luteum | Liu &al., 2012 |
| * Cryobacterium psychrophilum | Suzuku &al., 1997 |
| * Cryobacterium psychrotolerans | Zhang &al., 2007 |
| * Cryobacterium roopkundense | Reddy &al., 2010 |
| *** Cryobacterium sp. | Schuerger & Nicholson, 2016 |
| * Cytophaga aurantiaca | (Winogradsky, 1929) Reichenbach &al., 1989 |
| * Cytophaga hutchinsonii | Winogradsky, 1929 |
| * Cytophaga ssp. | Morita & Moyer, 2001 |
| † Deinococcus actinosclerus | Kim &al., 2018 |
| † Deinococcus aerius | Yang &al., 2009 |
| † Deinococcus aerolatus | Yoo &al., 2010 |
| † Deinococcus aerophilus | Yoo &al., 2010 |
| † Deinococcus aetherius | Yang &al., 2010 |
| † Deinococcus alpinitundrae | Callegan &al., 2008 |
| † Deinococcus altitudinis | Callegan &al., 2008 |
| † Deinococcus aluminii | Kim &al., 2018 |
| † Deinococcus antarcticus | ? |
| † Deinococcus apachensis | Rainey & da Costa, 2005 |
| † Deinococcus aquaticus | Im &al., 2008 |
| † Deinococcus aquatilis | Kampfer &al., 2008 |
| † Deinococcus aquivivus | Kaempferet &al., 2008 |
| † Deinococcus arenae | Lee &al., 2016 |
| † Deinococcus budaensis | Makk &al., 2016 |
| † Deinococcus caeni | Im &al., 2008 |
| † Deinococcus carri | ? |
| † Deinococcus callulosilyticus | Weon &al., 2007 |
| † Deinococcus citri | ? |
| † Deinococcus claudionis | Callegan &al., 2008 |
| † Deinococcus daejeonensis | Srinivasan &al., 2011 |
| † Deinococcus depolymerans | Asker &al., 2011 |
| † Deinococcus deserti | de Groot &al., 2005 |
| † Deinococcus erythromyxa | Brooks & Murray, 1981 |
| † Deinococcus ficus (also alkaliphile) | (Shashidhar & Bandekar, 2005) Kampfer, 2009 |
| *††† Deinococcus frigens | Hirsch &al., 2006 |
| † Deinococcus gammatolerans | Srinivasan &al., 2017 |
| † Deinococcus geothermalis (also a near-thermophile) | Ferreira &al., 1997 |
| † Deinococcus gobiensis | Yuan &al., 2009 |
| † Deinococcus grandis | Rainey &al., 1997 |
| † Deinococcus hibisci | Moya &al., 2018 |
| † Deinococcus hohokamensis | Rainey &al., 2005 |
| † Deinococcus hopiensis | Rainey &al., 2005 |
| † Deinococcus indicus | Suresh &al., 2004 |
| † Deinococcus knuensis | Lee &al., 2017 |
| † Deinococcus koreense | Kim &al., 2017 |
| † Deinococcus koreensis | Baek &al., 2018 |
| † Deinococcus lacus | Park &al., 2018 |
| † Deinococcus malanensis | Zhu &al., 2017 |
| † Deinococcus maricopensis | Rainey &al., 2005 |
| *† Deinococcus marmoris | Hirsch, 2004 |
| † Deinococcus metalli | ? |
| † Deinococcus metallilatus | ? |
| † Deinococcus misasensis | Asker &al., 2008 |
| † Deinococcus multiflagellatus | Kim &al., 2018 |
| † Deinococcus murrayi
(also a near-thermophile) |
Ferreira &al., 1997 |
| † Deinococcus navajonensis | Rainey &al., 2005 |
| † Deinococcus papagonensis | Rainey &al., 2005 |
| † Deinococcus peraridilitoris | Rainey &al., 2007 |
| † Deinococcus persicinus | Jeon &al., 2016 |
| † Deinococcus pimensis | Rainey &al., 2005 |
| † Deinococcus piscis | Shashidhar &al., 2009 |
| † Deinococcus planocerae | Lin &al., 2017 |
| † Deinococcus populi | Lee &al., 2014 |
| † Deinococcus proteolyticus | Kobatake &al., 1973? |
| † Deinococcus puniceus | ? |
| † Deinococcus radiodurans | Brooks & Murray, 1981 |
| † Deinococcus radiomollis | Callegan, 2008 |
| † Deinococcus radiophilus | Lewis, 1976 |
| † Deinococcus radiopugnans | Davis, 1963 |
| † Deinococcus radioresistens | Srinivasan &al., 2016 |
| † Deinococcus reticulitermitis | Chen &al., 2011 |
| † Deinococcus roseus | Asker &al., 2008 |
| † Deinococcus ruber | Kim &al., 2017 |
| † Deinococcus rufus | Wang &al., 2018 |
| † Deinococcus sahariens | Bouraoui &al., 2012 |
| † Deinococcus saudiensis | ? |
| *††† Deinococcus saxicola | Hirsch &al., 2006 |
| † Deinococcus sedimenti | Lee &al., 2017 |
| † Deinococcus seoulensis | ? |
| † Deinococcus soli | Zhang &al., 2011; Cha &al., 2016 |
| † Deinococcus sonorensis | Rainey &al., 205 |
| † Deinococcus taklimakanensis | Liu &al., 2017 |
| † Deinococcus wulumuqiensis | Wang &al., 2010 |
| † Deinococcus xibeiensis | Wang &al., 2010 |
| † Deinococcus xianganensis | Zheng &al., 2014 |
| † Deinococcus xinjangensis | Peng &al., 2009 |
| † Deinococcus yavapaiensis | Rainey &al., 2009 |
| † Deinococcus yunweiensis | Zhang &al., 2007 |
| ** Exiguobacterium sp. | Schuerger & Nicholson, 2016 |
| * Flavobacterium ssp. | Morita & Moyer, 2001 |
| * Gelidibacter ssp. | Morita & Moyer, 2001 |
| *+ Halomonas sp. ANT108
(also halotolerant) |
Wang &al., 2020 |
| * Hyphomonas adhaerens | Winer &al., 2000 |
| * Hyphomonas atlantica | Li &al., 2015 |
| * Hyphomonas beringensis | Li &al., 2015 |
| * Hyphomonas chukchiensis | Li &al., 2015 |
| * Hyphomonas hirschiana | Weiner &al., 1985 |
| * Hyphomonas jannaschiana | Weiner &al., 1985 |
| * Hyphomonas johnsonii | Weiner &al., 2000 |
| * Hyphomonas neptunium | (Leifson, 1964) Moore &al., 1984 |
| * Hyphomonas oceanitis | Weiner &al., 1985 |
| * Hyphomonas polymorpha | (Pongratz, 1957) Moore &al., 1984 |
| * Hyphomonas rosenbergii | Weiner &al., 2000 |
| * Hyphomonas sp. | Edwards &al., 2003 |
| * Methanococcoides ssp. | Morita & Moyer, 2001 |
| * Methanogenium ssp. | Morita & Moyer, 2001 |
| * Methanosarcina ssp. | Morita & Moyer, 2001 |
| * Methylococcus sp. | Morita & Moyer, 2001 |
| * Microbacterium ssp. | Morita & Moyer, 2001 |
| * Micrococcus ssp. | Morita & Moyer, 2001 |
| * Moritella ssp. | Morita & Moyer, 2001 |
| ***††† Natrialba magadii | (Tindall &al., 1984) Kamekura &al., 1997 |
| * Octadecabacter ssp. | Morita & Moyer, 2001 |
| ** Paenibacillus sp. | Schuerger & Nicholson, 2016 |
| * Phormidium ssp. | Morita & Moyer, 2001 |
| * Photobacterium ssp. | Morita & Moyer, 2001 |
| * Polaribacter ssp. | Morita & Moyer, 2001 |
| * Polaromonas ssp. | Morita & Moyer, 2001 |
| * Pseudomonas antarctica | Chauhan &al., 2015 |
| * Pseudomonas helmanticens | Saurabh &al., 2020 |
| * Pseudomonas meridiana | Chauhan &al., 2015 |
| * Pseudomonas proteolytica | Chauhan &al., 2015 |
| * Pseudomonas psychrophila | Yumoto &al., 2001 |
| * Pseudomonas ssp. | Morita & Moyer, 2001 |
| *† Psychrobacter adeliensis | Garcia-Lopez &al., 2014 |
| *† Psychrobacter aestuarii | Garcia-Lopez &al., 2014 |
| *† Psychrobacter alimentarius | Garcia-Lopez &al., 2014 |
| *† Psychrobacter aquaticus | Garcia-Lopez &al., 2014 |
| *† Psychrobacter aquimaris | Garcia-Lopez &al., 2014 |
| *† Psychrobacter arcticus | Garcia-Lopez &al., 2014 |
| *† Psychrobacter arenosus | Garcia-Lopez &al., 2014 |
| *† Psychrobacter celer | Garcia-Lopez &al., 2014 |
| *† Psychrobacter cibarius | Garcia-Lopez &al., 2014 |
| *† Psychrobacter cryohalolentis
(also a halophile?) |
Garcia-Lopez &al., 2014 |
| *† Psychrobacter faecalis | Garcia-Lopez &al., 2014 |
| *† Psychrobacter fozii | Garcia-Lopez &al., 2014 |
| *† Psychrobacter frigidicola | Garcia-Lopez &al., 2014 |
| *† Psychrobacter fulvigenes | Garcia-Lopez &al., 2014 |
| *† Psychrobacter glacincola | Garcia-Lopez &al., 2014 |
| *† Psychrobacter immobilis | Garcia-Lopez &al., 2014 |
| *† Psychrobacter jeotgali | Garcia-Lopez &al., 2014 |
| *† Psychrobacter luti | Garcia-Lopez &al., 2014 |
| *† Psychrobacter lutiphocae | Garcia-Lopez &al., 2014 |
| *† Psychrobacter marincola | Garcia-Lopez &al., 2014 |
| *† Psychrobacter maritimus | Garcia-Lopez &al., 2014 |
| *† Psychrobacter namhaensis | Garcia-Lopez &al., 2014 |
| *† Psychrobacter nivimaris | Garcia-Lopez &al., 2014 |
| *† Psychrobacter okhotskensis | Garcia-Lopez &al., 2014 |
| *† Psychrobacter pacificensis | Garcia-Lopez &al., 2014 |
| *† Psychrobacter phenylpyruvicus | Garcia-Lopez &al., 2014 |
| *† Psychrobacter piscatorii | Garcia-Lopez &al., 2014 |
| *† Psychrobacter proteolyticus | Garcia-Lopez &al., 2014 |
| *† Psychrobacter pulmonis | Garcia-Lopez &al., 2014 |
| *† Psychrobacter salsus | Garcia-Lopez &al., 2014 |
| *† Psychrobacter sanguinis | Garcia-Lopez &al., 2014 |
| *† Psychrobacter submarinus | Garcia-Lopez &al., 2014 |
| *† Psychrobacter urativorans | Garcia-Lopez &al., 2014 |
| *† Psychrobacter vallis | Garcia-Lopez &al., 2014 |
| * Psychroserpens ssp. | Morita & Moyer, 2001 |
| ** Rhodococcus sp. | Schuerger & Nicholson, 2016 |
| † Rubrobacter aplysinae | Kampfer &al., 2014 |
| † Rubrobacter bracarensis | Jurado &al., 2012 |
| † Rubrobacter calidifluminis | Albuquerque &al., 2014 |
| † Rubrobacter naiadicus | Albuquerque &al., 2014 |
| † Rubrobacter radiotolerans | Yoshinaka &al., 1973 |
| † Rubrobacter taiwanensis | Chen &al., 2004 |
| † Rubrobacter xylanophilus | Carreto &al., 1996 |
| † Rubrobacter wessexii | ? |
| * Shewanella arctica | Kim &al., 2012 |
| * Shewanella donghaensis | Yang &al., 2009 |
| * Shewanella frigidimarina | Bowman &al., 1997 |
| * Shewanella psychromarinicola | Hwang &al., 2019 |
| * Shewanella ssp. | Morita & Moyer, 2001 |
| ** Streptomyces sp. | Schuerger & Nicholson, 2016 |
| ** Serratia ficaria | Schuerger & Nicholson, 2016 |
| ** Serratia fonticola | Schuerger & Nicholson, 2016 |
| ** Serratia grimesii | Schuerger & Nicholson, 2016 |
| ** Serratia liquefaciens | Schuerger & Nicholson, 2016 |
| ** Serratia plymuthica | Schuerger & Nicholson, 2016 |
| ** Serratia quinivorans | Schuerger & Nicholson, 2016 |
| † Thermococcus gammatolerans | ? |
| * Vibrio ssp. | Morita & Moyer, 2001 |
References
Note: This section is still under construction!
Schuerger & Nicholson, 2016.
http://doi.org/10.1089/ast.2016.1587
Nicholson &al., 2013.
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